2.3 Plant Physiology: Photosynthesis

1. The Process and Importance

  • Definition: Conversion of light energy into chemical energy (glucose) from CO₂ and H₂O, releasing O₂.
  • Equation: 6CO₂ + 6H₂O + Light Energy → C₆H₁₂O₆ + 6O₂
  • Importance: Primary energy source, oxygen production, CO₂ regulation, fossil fuel formation.

2. Site of Photosynthesis: The Chloroplast

  • Location: Primarily in mesophyll cells of leaves.
  • Structure:
    • Double Membrane: Outer and Inner.
    • Stroma: Fluid-filled space; site of dark reactions (biosynthetic phase).
    • Thylakoids: Interconnected sacs; site of light reactions (photochemical phase).
    • Grana: Stacks of thylakoids.
    • Chlorophyll: Pigment embedded in thylakoid membranes.

3. Phases of Photosynthesis

  • 3.1. Light-Dependent Reactions (Photochemical Phase):
    • Site: Thylakoid membranes.
    • Process: Light energy activates chlorophyll, splits water (photolysis), transports electrons, produces ATP (photophosphorylation) and NADPH.
    • Summary: Light energy → Chemical energy (ATP, NADPH) + O₂.

3.2. Light-Independent Reactions (Biosynthetic Phase / Calvin Cycle)

  • Site: Stroma.
  • Process: Uses ATP and NADPH from light reactions.
    1. Carbon Fixation: CO₂ combines with RuBP (catalyzed by RuBisCO) to form PGA.
    2. Reduction: PGA converted to G3P using ATP and NADPH.
    3. Regeneration: RuBP regenerated from G3P using ATP.
  • Summary: CO₂ → Glucose (using ATP, NADPH).

4. Adaptations in Plants for Photosynthesis

  • Leaf Structure: Large surface area, thinness, cuticle, transparent epidermis, palisade/spongy mesophyll.
  • Stomata: Regulate CO₂ entry and water vapor release (opening/closing via K⁺ ion exchange).
  • Chloroplasts: Orient to maximize light absorption.
  • Vascular Bundles: Efficient transport of water and sugars.
  • Root System: Absorbs water and minerals.

5. Experiments to Show Necessity of Factors

  • Destarching: Keep plant in darkness to use up stored starch.
  • Starch Test (Iodine Test): Blue-black indicates starch.
  • Necessity of Light: Covered part of leaf (no light) shows no starch.
  • Necessity of CO₂: Plant in CO₂-free environment shows no starch.
  • Necessity of Chlorophyll: Non-green parts of variegated leaf show no starch.
  • Release of O₂: Aquatic plant in light releases bubbles that rekindle a glowing splint.

6. The Carbon Cycle

  • Definition: Biogeochemical cycle of carbon exchange among Earth's reservoirs.
  • Photosynthesis's Role: Removes atmospheric CO₂ and converts it into organic compounds, forming the base of food chains.
  • Key Processes: Photosynthesis, Consumption, Respiration, Decomposition, Fossil Fuel Formation, Combustion, Oceanic Carbon.