2.4 Plant Physiology: Chemical Coordination in Plants

1. Plant Growth Regulators (Phytohormones)

  • Chemical messengers regulating plant growth, development, and responses.

1.1. Auxins

  • Primary Role: Cell elongation and differentiation.
  • Physiological Effects:
    • Cell Elongation: Promotes stem growth.
    • Apical Dominance: Inhibits lateral bud growth.
    • Root Initiation: Promotes adventitious roots on cuttings.
    • Fruit Development: Promotes growth, prevents premature drop.
    • Herbicide Action: High concentrations act as selective herbicides (e.g., 2,4-D).

1.2. Gibberellins (GAs)

  • Primary Role: Stem elongation.
  • Physiological Effects:
    • Stem Elongation: Significant increase in length.
    • Seed Germination: Break dormancy, promote enzyme synthesis.
    • Flowering: Can induce flowering in some plants.
    • Fruit Development: Promote fruit growth (e.g., grapes).
    • Bolting: Stem elongation in rosette plants.

1.3. Cytokinins

  • Primary Role: Cell division and differentiation.
  • Physiological Effects:
    • Cell Division: Promotes cell proliferation.
    • Delay Senescence: Delays aging and yellowing of leaves.
    • Break Apical Dominance: Promotes lateral bud growth.
    • Chloroplast Development: Promotes chloroplast formation.

1.4. Abscisic Acid (ABA)

  • Primary Role: Stress hormone, growth inhibitor.
  • Physiological Effects:
    • Stomatal Closure: Reduces water loss during stress.
    • Seed Dormancy: Induces and maintains dormancy.
    • Bud Dormancy: Promotes dormancy in buds.
    • Abscission: Promotes shedding of leaves, fruits, flowers.

1.5. Ethylene

  • Primary Role: Fruit ripening and senescence (gaseous hormone).
  • Physiological Effects:
    • Fruit Ripening: Promotes ripening of climacteric fruits.
    • Senescence and Abscission: Accelerates aging and shedding.
    • Triple Response: Inhibition of stem elongation, radial swelling, horizontal growth in seedlings.

2. Tropic Movements in Plants

  • Directional growth responses to external stimuli, regulated by hormones (especially auxins).

2.1. Phototropism

  • Definition: Growth movement in response to light.
  • Mechanism: Auxins migrate to shaded side, causing differential cell elongation.
  • Types:
    • Positive: Towards light (shoots).
    • Negative: Away from light (roots, weak).

2.2. Geotropism (Gravitropism)

  • Definition: Growth movement in response to gravity.
  • Mechanism: Auxins accumulate on the lower side; roots are more sensitive to auxin inhibition.
  • Types:
    • Positive: Towards gravity (roots).
    • Negative: Away from gravity (shoots).

2.3. Hydrotropism

  • Definition: Growth movement in response to water.
  • Type: Roots typically exhibit positive hydrotropism (grow towards water).

2.4. Thigmotropism

  • Definition: Growth movement in response to touch or physical contact.
  • Mechanism: Differential growth causes coiling around support.
  • Example: Coiling of tendrils.

2.5. Chemotropism

  • Definition: Growth movement in response to a chemical stimulus.
  • Example: Pollen tube growth towards the ovule (positive chemotropism).