Unit 1: Reproduction

Chapter 1.1: Sexual Reproduction in Flowering Plants

Flower Structure

  • Definition: The reproductive organ of angiosperms.
  • Main Parts:
    • Calyx (Sepals): Protects the bud.
    • Corolla (Petals): Attracts pollinators.
    • Androecium (Stamens): Male reproductive part.
    • Gynoecium (Pistil/Carpel): Female reproductive part.

Androecium (Male)

  • Composed of stamens.
  • Each stamen has:
    • Filament: The stalk.
    • Anther: Produces pollen.

Gynoecium (Female)

  • Composed of carpels.
  • Each carpel has:
    • Ovary: Contains ovules.
    • Style: Connects ovary to stigma.
    • Stigma: Receptive tip for pollen.

Male Gametophyte

  • Microsporangium (Pollen Sac):
    • Located in the anther.
    • Contains sporogenous tissue.
  • Microsporogenesis:
    • Formation of microspores from microspore mother cells (MMCs) via meiosis.
  • Pollen Grain:
    • Represents the male gametophyte.

Pollen Grain Structure

  • Exine: Hard outer layer (sporopollenin).
  • Intine: Thin inner layer (pectocellulose).
  • Two Cells:
    • Vegetative Cell: Forms pollen tube.
    • Generative Cell: Forms two male gametes.

Female Gametophyte

  • Ovule (Megasporangium):
    • Develops into a seed after fertilization.
    • Anatropous Ovule: Most common, inverted type.
  • Megasporogenesis:
    • Formation of megaspores from a megaspore mother cell (MMC) via meiosis.
  • Embryo Sac:
    • The female gametophyte.

Embryo Sac Development

  • Develops from a single functional megaspore.
  • 7-celled, 8-nucleate structure:
    • Egg apparatus: 1 egg cell, 2 synergids.
    • Central cell: 2 polar nuclei.
    • Antipodal cells: 3 cells at the chalazal end.

Pollination

  • Definition: Transfer of pollen from anther to stigma.
  • Types:
    • Autogamy: Self-pollination (same flower).
    • Geitonogamy: Same plant, different flower.
    • Xenogamy: Cross-pollination (different plant).

Agents of Pollination

  • Abiotic:
    • Wind (Anemophily): Grasses, corn.
    • Water (Hydrophily): Vallisneria, Zostera.
  • Biotic:
    • Insects (Entomophily): Bees, butterflies.
    • Animals (Zoophily): Birds, bats.

Outbreeding Devices

  • Mechanisms to encourage cross-pollination.
    • Dichogamy: Anthers and stigma mature at different times.
    • Herkogamy: Physical barrier between anther and stigma.
    • Heterostyly: Different lengths of styles and stamens.
    • Self-incompatibility: Genetic mechanism to prevent self-pollen germination.
    • Unisexuality: Male and female flowers on different plants.

Pollen-Pistil Interaction

  • The sequence of events from pollen deposition to pollen tube entry into the ovule.
  • Recognition: Pistil recognizes compatible pollen.
  • Germination: Pollen tube grows.
  • Entry into Ovule: Pollen tube enters the ovule, usually through the micropyle.

Artificial Hybridization

  • A crop improvement program.
  • Steps:
    1. Emasculation: Removal of anthers.
    2. Bagging: Covering the flower to prevent unwanted pollination.
    3. Pollination: Dusting with desired pollen.
    4. Rebagging: Until fruit develops.

Double Fertilization

  • A unique event in flowering plants.
  • Syngamy: One male gamete + egg cell -> Zygote (2n).
  • Triple Fusion: Other male gamete + two polar nuclei -> Primary Endosperm Nucleus (PEN) (3n).

Post-Fertilization Events

  • Ovary -> Fruit
  • Ovules -> Seeds
  • Zygote -> Embryo
  • PEN -> Endosperm

Seed and Fruit

  • Seed:
    • Albuminous: Retains endosperm (e.g., wheat, maize).
    • Non-Albuminous: No residual endosperm (e.g., pea, groundnut).
  • Fruit:
    • True Fruit: Develops only from the ovary.
    • False Fruit: Develops from the ovary and other floral parts.

Special Modes of Reproduction

  • Apomixis: Seed formation without fertilization.
  • Parthenocarpy: Fruit development without fertilization (seedless fruits).
  • Polyembryony: Occurrence of more than one embryo in a seed.

Seed and Fruit Dispersal

  • Significance:
    • Prevents overcrowding.
    • Colonization of new areas.
    • Promotes genetic diversity.
  • Agents:
    • Wind, water, animals, explosive mechanisms.