Unit 2: Genetics and Evolution

Chapter 2: Molecular Basis of Inheritance

Search for Genetic Material

  • Griffith's Experiment: Showed a "transforming principle" could change bacteria.
  • Avery, MacLeod, McCarty: Identified DNA as the transforming principle.
  • Hershey-Chase Experiment: Confirmed DNA is the genetic material using bacteriophages.

DNA Structure

  • Watson & Crick Model: Double helix with two anti-parallel strands.
  • Backbone: Sugar-phosphate.
  • Base Pairing:
    • Adenine (A) with Thymine (T) (2 H-bonds)
    • Guanine (G) with Cytosine (C) (3 H-bonds)
  • Chargaff’s Rule: A=T and G=C.

DNA Packaging

  • Nucleosome: DNA wrapped around a core of 8 histone proteins.
  • Chromatin: A chain of nucleosomes ("beads-on-string").
  • Chromosomes: Highly condensed chromatin.

Central Dogma

  • Describes the flow of genetic information:
    DNA → RNA → Protein
  • Replication: DNA → DNA
  • Transcription: DNA → RNA
  • Translation: RNA → Protein
  • Reverse Transcription: RNA → DNA (in retroviruses).

DNA Replication

  • Semi-conservative: Each new DNA has one old and one new strand.
  • Proven by: Meselson and Stahl experiment.
  • Key Enzymes:
    • Helicase: Unwinds DNA.
    • DNA Polymerase: Synthesizes new DNA.
    • Ligase: Joins DNA fragments.

Transcription

  • Definition: Synthesis of RNA from a DNA template.
  • In Eukaryotes:
    • Occurs in the nucleus.
    • Produces hnRNA (pre-mRNA).
  • Post-transcriptional Processing:
    • Splicing: Removes introns, joins exons.
    • Capping: Adds a cap to the 5' end.
    • Tailing: Adds a poly-A tail to the 3' end.

Genetic Code

  • Triplet Code: 3 bases = 1 codon.
  • Degenerate: Multiple codons for one amino acid.
  • Unambiguous: One codon for one amino acid.
  • Universal: Same code in most organisms.
  • Start Codon: AUG (Methionine).
  • Stop Codons: UAA, UAG, UGA.

Translation

  • Definition: Synthesis of protein from an mRNA template.
  • Occurs on: Ribosomes.
  • Steps:
    1. Initiation: Ribosome binds to mRNA.
    2. Elongation: Polypeptide chain grows.
    3. Termination: Stop codon is reached.

Gene Expression: Lac Operon

  • An inducible operon in E. coli for lactose metabolism.
  • Lactose Absent: Repressor binds to operator, operon is OFF.
  • Lactose Present: Lactose (inducer) binds to repressor, operon is ON.

Human Genome Project (HGP)

  • Goal: To sequence the entire human genome.
  • Key Findings:
    • ~3 billion base pairs.
    • ~20,000-25,000 genes.
  • Applications: Disease diagnosis, gene therapy, forensic science.

DNA Fingerprinting

  • Principle: Based on Variable Number Tandem Repeats (VNTRs).
  • Technique: Involves restriction digestion, gel electrophoresis, and hybridization with a VNTR probe.
  • Applications:
    • Forensic science (crime investigation).
    • Paternity testing.
    • Conservation biology.